Reverse Coiled Gastropods

Land & Freshwater Gastropods

Although not as well-publicized as their marine gastropod relatives, the shells of certain few snail species inhabiting the land and fresh waters are known to occur in the abnormally-reversed-coiled condition. Actually, the number of species and individual specimens with reversed symmetry appears to be greater among this group (with terrestrial pulmonate shells accounting for the lion's share) than among seashells - rather surprising considering that far fewer species and individuals of nonmarine than of marine shells have come to our attention.

Aquatic Prosobranchs

Ampulariidae Pila ampullacea (Linnaeus, 1758) 5.86 Cavite, Luzon, Philippines. (1994).
Ampulariidae Pila globosa (Swainson, 1821) 3.41 Jelatura, Sri Lanka. 1981.
Viviparidae Campeloma decisum (Say, 1817) 31.0 Des Plaines R., Illinois, USA. (1986).
Viviparidae Campeloma lima (Anthony, 1860) 2.58 Doctortown, Georgia, USA. 1974.
Viviparidae Viviparus viviparus (Millet, 1813) 1.83 Overrijssel, Zwartsluis, Zwartewater, Netherlands. (2003).
Nassariidae Nassodonta dorri (Wattebled, 1886) 1.31 Pthan Ri, Vietnam. 2005.

Aquatic Pulmonate

Lymnaeidae Lymnaea stagnalis (Linnaeus, 1758) 2.7 Paks, Danubevally, Hungary. 1983.

Terrestrial Prosobranchs

Diplommatinidae Palaina taeniolata hyalina Quadras and Möllendorff, 1894 0.23 Agana, Guam. (1986). (species is normally sinistral) +
Pupinidae Schistoloma alta sibuyanica Bartsch, 1915 2.55 Sibuyan Is. Philippines. 1940. (species is normally dextral)

Terrestrial Pulmonates

Acavidae Acavus phoenix (L. Pfeiffer, 1854) 5.51 Horana, Sri Lanka. (1988).
Acavidae Acavus haemastoma (Linnaeus, 1758) 3.98 Galle, Sri Lanka. 1988.
Achatinidae Achatina fulica (Bowdich, 1822) 8.07 Madagascar. (1988). +
Achatinidae Achatina immaculata Lamarck, 1822 8.78 Curepipe, Mauritius. (1973).
Achatinidae Achatina zebra (Bruguière, 1792) 6.50 near Jeffreys Bay, South Africa. (2006).
Achatinidae Archachatina ventricosa (Gould, 1850) 7.45 Grand Bassam, Ivory Coast. (1995).
Clausiliidae Alinda biplicata (Montagu, 1803) 1.40 Budapest, Hungary. 1983. (species is normally sinistral)
Clausiliidae Cochlodina laminata (Montagu, 1803) 1.60 Oxon, England. (1971). (species is normally sinistral)
Clausiliidae Leucostigma candidescens (Rossmässler, 1836) 1.90 Avezzanno, Abruzzi Prov., Italy 2001. (species is normally sinistral)
Ariophantidae Elaphroconcha javacensis (Férussac, 1821) 2.10 Tupang, Java. 2000.
Bradybaenidae Calocochlia pan (Broderip, 1841) 5.05 Bohol, Philippines. (1994). +
Bradybaenidae Helicostyla daphnis (Broderip, 1841) 4.55 Cebu, Philippines. 1999.
Bradybaenidae Helicostyla effusa (L. Pfeiffer, 1842) 4.82 Batangas, Philippines. (1999).
Bradybaenidae Helicostyla pithogaster (Férussac, 1821) 5.05 Luzon, Philippines. (1997).
Bulimulidae Corona perversa (Swainson, 1820) 5.75 Serra de Navio, Amapá, Brasil. 2002. (species is normally sinistral)
Bulimulidae Corona regalis (Hupé, 1857) 6.98 Meruri, Mato Grosso, Brasil. 1980. (species is normally sinistral)
Bulimulidae Liguus fasciatus (Müller, 1774) Pincrest Hammock no. 1, Florida, USA. 1940.
Bulimulidae Liguus virgineus (Linnaeus, 1758) 3.70 Haiti. (1975).
Bulimulidae Rabdotus mooreanus (L. Pfeiffer, 1868) 2.16 Ft. Worth, Texas, USA. 1977.
Bulimulidae Thaumastus largillierti (Philippi, 1848) 5.57 Arraial do Cabo, Brasil. 2001.
Camaenidae Amphidromus chloris (Reeve, 1848) 3.70 Philippines. (1986). (species is normally sinistral)
Camaenidae Amphidromus latestrigatus Schepman, 1892 3.60 small island near Sawu Island, East Indonesia. (2006). (species is normally sinistral)
Camaenidae Caracolus marginella (Gmelin, 1791) 3.94 Wainwright Hammock, Miami, Florida, USA. 2000.
Camaenidae Obba moricandi (L. Pfeiffer, 1842) 2.68 Samar, Philippines. (1995).
Discidae Anguispira alternata (Say, 1817) 2.02 Medina, Michigan, USA. 1941.
Discidae Anguispira strongylodes (L. Pfeiffer, 1854) 1.40 Marianna, Florida, USA. 1990.
Helicarionidae Ryssota otaheitana (Férussac, 1821) 7.63 Bohol, Philippines. (1997).
Zonitidae Ventridens cerinoideus (Anthony, 1865) 0.62 Jacksonville, Florida, USA. 2005.
Helicidae Cornu aspersum (Müller, 1774) 2.76 Newark, England. 1978.
Helicidae Eobania vermiculata (Müller, 1774) 3.56 Languedoc, France. (1991).
Helicidae Helix lucorum Linnaeus, 1758 4.62 Southern Greece. (1993).
Helicidae Helix pomatia Linnaeus, 1758 4.26 Geneva, Switzerland. (1974).
Helicidae Otala punctata (Müller, 1774) 4.09 Tarragona, Spain (2000).
Megalobulimidae Megalobulimus chianostoma (Mørch, 1852) 11.18 Arraial do Cabo, Brasil. 2001.
Polygyridae Daedalochila avara (Say, 1818) 0.66 Jacksonville, Florida, USA. 1975.
Polygyridae Polygyra septemvolva Say, 1818 0.73 Levy Co., Florida, USA. 2005.
Polygyridae Praticolella species 0.50 El Limon, Mexico. 1989.
Polygyridae Xolotrema fosteri (F. C. Baker, 1921) 1.53 St. Louis, Missouri, USA. 1969.
Pupillidae Vertigo rugosula Sterki, 1890 0.22 Jacksonville, Florida, USA. 1975.
Ceriidae Cerion incanum (A. Binney, 1851) 1.87 Monroe Co., Florida, USA. 1999.
Spiraxidae Euglandina liebmanni (L. Pfeiffer, 1846) 2.95 Guerrero, Mexico. (1992).

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